¹University Medical Center Hamburg-Eppendorf, Laboratory for Radiobiology and Experimental Radiooncology, Hamburg, Germany; ²University Medical Center Hamburg-Eppendorf, Department of Radiotherapy and Radiation Oncology, Hamburg, Germany; ³German Cancer Consortium, Partner Site Tuebingen and German Cancer Research Center, Heidelberg, Germany; ⁴University of Tuebingen, Division of Radiobiology & Molecular Environmental Research, Department of Radiation Oncology, Tuebingen, Germany; ⁵Technische Universität Dresden, National Center for Radiation Research in Oncology, Molecular and Cellular Radiobiology, Dresden, Germany; ⁶Helmholtz-Zentrum Dresden-Rossendorf, Institute of Radiooncology-OncoRay, Dresden, Germany; ⁷German Cancer Consortium, Partner Site Dresden: German Cancer Research Center, Heidelberg, Germany; ⁸University Hospital Carl Gustav Carus, Technische Universität Dresden, Department of Radiotherapy and Radiation Oncology, Dresden, Germany

(1)Institute of Nuclear Physics PAN, Proton Radiotherapy Group, Krakow, Poland;(2)Loma Linda University, Department of Basic Sciences, Loma Linda, USA;(3)Northwestern Medicine Chicago Proton Center, Medical Physics and Dosimetry, Warrenville, USA;(4)Northern Illinois University, Department of Physics, DeKalb, USA;(5)Santa Cruz Institute for Particle Physics- University of California, Physics Department, Santa Cruz, USA;(6)University of California, Radiation Oncology, San Francisco, USA;

¹Brown University, Radiation Oncology, Providence, USA